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Samanosuke and Jacques can gain elemental weapons, changing their fighting style and allowing access to magical attacks. Defeating enemies reaps souls, which when absorbed can restore health and magic, be used as currency for purchasing ammunition and health items, and allow Samanosuke or Jacques to enter a more powerful Onimusha state for a short time.1 If the player collects certain items during the game and completes it, Samanosuke will be able to use his weapons from the first game in the next playthrough. Armors with different properties can be obtained in the game but they require some items in order to work.

During certain portions of the story, players also get to control Michelle, who relies on firearms. Another minigame gives the players control over the lance fighter Heihachiro. Unlike Samanosuke and Jacques, Heihachiro and Michelle do not possess upgradeable weapons and cannot enter the Oni mode.

New to this game is a focus on time travel when solving puzzles. For example, if Samanosuke comes across a door in the present that has become too withered to open, Jacques will need to open the door in the past so that it will stay open for Samanosuke to progress. Things Samanosuke does in the present will not affect things in the past.3 However, Ako is able to transfer some items between both timelines.

The game opens with the master swordsman Samanosuke Akechi destroying the monstrous Genma army in Japan 1582 and goes to Honn?-ji Temple to beat his nemesis, Nobunaga Oda. Samanosuke defeats the warrior Mori Ranmaru and approaches Nobunaga. However, before they can fight, a portal opens in the floor and Samanosuke is pulled through it. It then jumps to modern-day Paris 2004, featuring a man named Jacques Blanc fighting the Genma army then begins its invasion of Paris near the Arc de Triomphe. Samanosuke appears from a portal in Paris and rescues Jacques from the Genma. The time portal reappears and pulls Jacques and Philippe through to feudal Japan. Samanosuke then meets Michelle, Jacques girlfriend, near the Arc and makes his way towards the roof. Across his fights, Samanosuke briefly meets his old enemy, the scientist Guildenstern, who is responsible for sending him and the Genma to the future.

Jacques Blanc and Philippe arrive in 16th century Japan. However, Philippe dies of mortal wounds from the previous battle after the time warp. Upon arriving, an Oni spirit grants him powers in order to assist this timelines Samanosuke on his way to Honn?-ji Temple. The Oni also gives Jacques a spirit named Ako to assist him in his quest. To return to their own time period, Samanosuke and Jacques must save the respective timelines they are in from destruction. The Samanosuke in 2004 Paris teams up with Michelle and Jacques son Henri to save the city from destruction while Jacques and the pasts Samanosuke struggle to fight Nobunaga in feudal Japan. Across his journey, Jacques and the pasts Samanosuke meet Tadakatsu Heihachir? Honda who is linked with the Oda clan. However, Heihachir? joins decides to join Samanosuke and Jacques cause. The future Samanosuke also encounters the futures Ranmaru who has become a Genma warrior.

Following several battles, Samanosuke manages to defeat the Genma forces from the future by defeating Guildenstern and Ranmaru. Heihachir? is killed by the pasts Ranmaru and Jacques avenges him when reaching Honn?-ji Temple. Although Nobunaga appears to be defeated and Jacques and the future Samanosuke start switching places, Nobunaga recovers and kills the pasts Samanosuke. Jacques reunites with his family but Henri is attacked by the futures Ranmaru. However, Jacques Oni powers are transferred to Henris body, saving the boys life. Back in the past, Samanosuke absorbs his alternate self murdered by Nobunaga to increase his powers and engage in a final fight. Samanosuke emerges victorious and absorbs Nobunagas soul into his Oni Gauntlet to stop him from reviving again. He then starts a journey with Ako to seal his Gauntlet.

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